The other great group of seed plants is known as the Angiosperms, and to it belong all those seed plants which are most commonly met in this region. The distribution of Angiosperms is a very much more difficult question than that of Gymnosperms; for while there are only about four hundred kinds of living Gymnosperms, there are more than one hundred thousand kinds of living Angiosperms. In presenting the distribution of this great group, it will be necessary to consider its two main divisions separately, for they differ from one another very much. One of the groups is known as the Monocotyledons, to which belong such forms as the grasses, lilies, palms, orchids, etc.

Some prominent facts in reference to the geographical distribution of these Monocotyledons are as follows: They contain four great families, which include almost one-half of their number, and which have become world-wide in their distribution. These families are the grasses, the sedges, the lilies, and the irises. This world-wide distribution means that these families have succeeded in adapting themselves to every condition of soil and climate. In this world distribution the grasses easily lead, not only among Monocotyledons, but among all seed plants.

Another fact in reference to the Monocotyledons is that they include an unusual number of families which are entirely aquatic in their habit. These aquatic families are also world-wide in their distribution, so far as fresh and brackish waters can be called world-wide. It is important to notice that while the world families which belong to the land have worked out about ten thousand different forms, the world families which belong to the water have worked out considerably less than two hundred different forms. This seems to indicate that the great number in the one case is due to the very diverse conditions of the land, while the small number in the latter case is due to the very uniform conditions of water life.

A third fact of importance is that the Monocotyledons are mainly massed in the tropics, and in this sense are almost an exact contrast to the Conifers we have been considering above. The same effect of separation in working out diversity in structure is shown by the Monocotyledons as was shown by the eastern and western Cycads, and the northern and southern Conifers. For example, the palms represent the great tree group of Monocotyledons, and are restricted to the tropics as rigidly as are the Cycads. They are found in about equal numbers in the eastern and western tropics, but there are no forms in common. The eastern and western forms have become so different that they might almost be regarded as different families.

The Monocotyledons are also somewhat famous for the number of air plants which they contain, that is, plants which have sometimes been called "perchers," because they fasten themselves upon trunks and branches and supports of various kinds, and absorb what they need directly from the air. It is a notable fact that these so-called "perchers" are very much more abundant in the western tropics than in the eastern. An explanation for this is to be found in the fact that the western tropics have a very much greater rainfall; in fact, in the rainy woods of the Amazon region the air is saturated with water, and everything is dripping.

One of the facts in connection with the distribution of Monocotyledons is quite puzzling, and that is the very poor representation of the whole group in the southern hemisphere. In examining the distribution of other groups in the southern hemisphere, it is found that Australia and its general vicinity is prolific in peculiar forms. In the case of the Monocotyledons, however, the Australasian region is the most poverty-stricken one in all the southern hemisphere just why the southern hemisphere in general, and the Australasian region in particular, are unfavorable for Monocotyledons, it is hard to say. Of course in these cases the world families already mentioned are represented.