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While fishes were numerous and large in the Devonian, throughout the Carboniferous they began to decline. By this time the land area had much increased, land plants became very abundant, there were immense forests of tropical vegetation, great swamps and peat bogs all of which later sank below sea level became covered up and changed into coal. Immense lizards lived in these forests and along the sea shores; these were the first land animals. At the close of the Carboniferous great changes took place; greater changes than at any time since the close of the Archaean. So marked were the changes at this time that it marks a new era in the geological history of the earth. All preceding the close of the Carboniferous is regarded as ancient geology; all since then as modern geology. It was at this time that plants and animals were represented by new forms more like those now living. The geological age following the Carboniferous is the Triassic. With this age began our modern sharks and fishes. They did not become abundant until the Jurassic and Cretaceous. All of the earlier sharks had strong spines in front of each dorsal fin and broad teeth made for crushing. One form of these known as Cestracionts were very abundant till the end of the Cretaceous. In the early Triassic they began to decline and the sharks, with pointed teeth, increased. These sharks, with pointed teeth, but rounded on the edges, commenced back in the Carboniferous. During the Triassic the sharks, with lancet shaped teeth, such as are now possessed by nearly all our sharks, commenced in small numbers. One of the important differences between the ganoids and the teleosts or true fishes is in the tail vertebrae. In the ganoids the tail vertebrae decrease gradually in size and curve upwards in the upper lobe of the tail. In the teleosts the tail vertebrae ends a short distance in front of ends of the middle fin rays of the tail fin. In the ganoids the upper lobe of the tail fin is the largest. In the teleosts both lobes are nearly the same size. |
The tail of the ganoid fish is called heterocercal, that of our modern or teleost fishes is homocercal. The tail of all early ganoids was strongly heterocercal. In the Triassic and Jurassic its lobes in many cases became nearly equal, approaching the homocercal tail. The tails of all sharks are heterocercal, of all modern fishes it is homocercal except in a few families, as the cod and related fishes, it is Isocercal; that is, the vertebrae decrease in size, but do not form an upward curve. So far as we know the Shad family is the first of our teleosts or true fishes to appear, and these were quite abundant in the early part of the Triassic. |
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